To add to above how exactly zipalign works -
In an Android operating environment, data files stored in each application package are accessed by multiple processes, for example, the installer will read the data manifest to determine the associated permissions; the system server can read these resources for multiple reasons, like displaying notifications; the Home application, for example, will read resources to get the application’s name and icon. Since Android is based on a a true multi-tasking operating infrastructure, these files are continually and repeatedly accessed. Finally, but not least, the application itself reads the manifest data.
As Android is Linux-based, memory-mapping plays a key role in efficient handling of processes. Essentially, the optimal alignment for the Android OS’ resource-handling code is 4-byte boundaries. What this means is that, if APKs are memory-mapped to 4-byte boundaries, and aligned accordingly, the OS will not need to ‘read through’ the whole application package to get to the desired data manifest. Every system process will know in advance where to look for it’s desired resources, and hence will execute much smoother and faster.
Summing it up, zipaligning an APK results in all uncompressed data within the package to be aligned on 4-byte boundaries, allowing all portions to be accessed directly with the memory-map. RAM consumption is lowered while execution because the querying code doesn’t have to read through the entire application package.